Jenny,

What is the average distance between mature trees in the forest?  You write:  
"I know it depends on the kind of forest and what kind of disturbance there has 
been,"  But that is the entire point. The answer is dependant on a variety of 
factors, so that a single answer is not possible.  We have had similar 
discussions on the average height of trees.  Look at how the trees grow.  
Initially a patch of seedlings grow very close together with several seedling 
per square foot.  Over time, some of the seedlings die out and the average 
spacing increases between the stems.  The spacing is greater in saplings. It 
increases on maturity.  As trees begin to die in a mature forest the spacing 
between the stems continues to increase.  It increases through time for a 
particular species.  Beyond the initial rapid thinning phase in which there are 
many trees already growing to fill the space lost by a dead tree, there is a 
point at which the loss of a tree will open up a space in the canopy and forest 
floor that will be filled by new growth.  This is where the period of 
disturbance comes into play.  The shorter the period of small disturbance the 
more common these denser new growth patches will be in the forest.  The size of 
the disturbance likewise plays a role in determining the size of the openings 
and the size of the denser new growth patches.  There is the size of the 
species in the mix.  Some trees simply grow bigger than others.  Those species 
that grow larger, will support a larger canopy, and have a greater trunk 
spacing than small species in areas that are relatively undisturbed.  A 
patchwork of mixed species may have a denser packing than those of a single or 
limited number of species.  The species present will also impact the number and 
type of trees in the understory.  

So there may be measurements made of the spacing in a particular forest, but 
that does not really answer the question.  It is a snapshot in time.  It tells 
you what the density of stems is in this particular forest, at this particular 
point in time, with this particular mix of species, and this particular 
disturbance history. Numbers can be generated, but are they meaningful?  A 
density measurement taken earlier in time, or later in time will give you 
different results.  The measurement is dynamic and changes with time, so that a 
simple answer that can be applied broadly can't really be found.   What I 
suggest doing is to try to reconsider what exactly you are trying to determine, 
what are you trying to accomplish, and find a different way to approach the 
problem.

Ed Frank  

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