Ed:

   I did a little on-line research and came across one interesting
article that I could understand to allow me, or reminded me, that I
could come at this logging effect on white pine genetics from a
different angle, one I am embarrassed that I didn't think of before.
It actually ties into something I learned from reading about
provenance trials of Norway spruce years ago, and a conversation I had
with a person in the UK who is very familiar with Norway spruce
populations.

   When I wrote my post about the idea that selective logging (maybe
two or three times is what I had in mind) could have affected the
genetic make up of the subsequent forest, I was of course thinking of
a specific place. this could cause some genetic drift, but as you
point out, not very substantial.

   What I am reminded of now is that different areas in the same area
rather close to each other can have white pines of substantially
different genetic make-up. I came across an article that talked about
this in Ontario.  One conclusion from this article was a bit like what
I reported before about the Clover Run pines in WV--that is that when
selecting seed to plant in a given area, it is not necessarily the
seed from the same area that would be best.  There is a very dramatic
example of this in Scandinavia with Norway spruce, caused by having
the area being repopulated after the most recent ice advance by Norway
spruce migrating from two different directions. Of course after the
most recent glaciations in North America, an uneven retreat of the ice
may have resulted in an uneven repopulation by white pine of various
areas.

   But in a way the study (which I admit I read only the abstract for)
is surprising, given that white pine pollen is spread by the wind,
which might suggest that populations over a wider area could be more
homogeneous.  With Norway spruce, a provenance trial done by Dr. John
Genys many years ago seemed to suggest something similar with Norway
spruce.  Seed collected from areas near each other produced clearly
different results.  Of  course,  the seed could have been collected in
different ways (which I won’t go into now), which perhaps could
account for the differences. Also, the provenances of Norway spruce in
Europe are somewhat muddied by having NS planted from seed collected
in one area and then planted in another for hundreds of years.

   Anyway, here is what I think MAY be a possibility here.  In the
past there were specific areas that had especially tall white
pines—tall because of a “superior” genetic strain, superior even to
those growing in areas not that far away.  Then as
civilization/logging came, those populations were not selectively
logged, but these stands of especially tall trees—and their specific
collection/arrangement of genes—were entirely wiped out.  All the
pines were cut, and the land in that area did not grow back with white
pine trees for one reason or another. There may have been some
"superior" strains of white pine that simply became "extinct."

   Of course what I am talking about here would not be an effect  from
specificlly targeting areas with the tallest pines (as with
specifically targeting the best trees to cut in a stand) and then
cutting the pines and converting the areas to other uses.  The process
would be rather random, except to the degree that the best stands
would be the most attractive to cut first, whether on the best sites
or not.  Of course, the richest sites would naturally have been
converted to farming.

   From what you understand, can this make sense?

   Of course, although I am trying to support the idea that white
pines taller than what we now see could have once existed, I am still
“skeptical” about 250 foot white pines.  I am just trying to explore
all possibilities, all possible explanations.

   --Gaines McMartin
   ---------------------------------------------------------

On 1/6/10, Edward Frank <[email protected]> wrote:
> Gaines,
>
> Basically, No you are incorrect.  In order to change the genetic footprint
> of the species you would need to cut only those superior trees, kill all of
> their offspring, and not cut any of the poorer quality trees, over the
> course of tens of generations.  Essentially all of the big trees in a stand
> were cut, so there was no selection.  The genetics of the big trees having
> been there for hundreds of years would be represented in the smaller trees
> that were too small to cut.  Even after the initial trees were removed,
> those trees with superior genetics would tend to out-compete those that were
> the offspring of poorer trees and would dominate the second growth forest.
> If you did this a hundred times you would not get a genetic drift, .unless
> you were targeting a specific genetic definable trait while at the same time
> leaving all of the other competing traits intact,  People have suggested
> this before, and the idea is simply wrong.  You can't change the genetic
> footprint by cutting all of the big pines, and it is unlikely that targeted
> high grading would even have a noticeable effect without dozens of
> generations of repetition.  There are specific environmental effects that
> can cause a genetic drift, but not generic logging operations.
>
> Ed Frank
>

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