Holy Mackerel! I can't believe the way you guys keep this discussion going (and 
others too)! Besides that, I can't believe how much you guys write in each of 
your responses. How can you come with so much to say?
(Hey, I'm not knocking it. I just can't believe it.)
By the way, this response doesn't need to be archived on the site. I just had 
to say this. This discussion is like the Energizer Bunny, if you know what I 
mean.

--- On Thu, 1/7/10, Gaines McMartin <[email protected]> wrote:


From: Gaines McMartin <[email protected]>
Subject: Re: [ENTS] Re: White pine growth rates--something of interest about 
growth possibilities
To: [email protected]
Cc: [email protected]
Date: Thursday, January 7, 2010, 8:40 AM


Ed:

   All the environmental factors causing one pine tree--or group of
pines--to grow taller than others I pretty much understand.  As for
the isolated populations:  I am not sure that is what is in play in
either Ontario with white pine, or in Europe with Norway spruce.  The
disparate results that popped up in Dr. Genys's trials were from
generally forested areas, not isolated areas.  And, unless I
misunderstood the article about white pine in Ontario, whose summary I
read, population isolation was not the issue in the variations in
white pine in Ontario. I commented in that post that I was surprized
that pollen drift has not eliminated that kind of variation, but
apparently it has not.

   Maybe one more thought about provenance trials--or their
interpretation.  As you pointed out, after selective logging in which
the best trees are cut, even if two or three times, and even if some
selectiing out of the best genes has occurred, the resultant
population will normally have a large enough number of trees so that
they will have to compete, and thus the individuals having the best
"growth" genes will dominate again, restoring the previous balance.

   In the same vein, provenance trials often have a short duration,
and the results after 3 or 4 years are results before any competition
occurs between individuals in the population being studied.  And
sometimes--although from my observations, not usually--the best
individuals after 4 years a may not be the best over a longer period
of time. Trees in a mature white pine stand and a mature Norway spruce
stand will be spaced an average of at least 25 feet apart.  I won't
try to do any precise calculations, but these trees will be a very
small fraction of the initial population.  So, to get a beautiful
stand of fast growing "superior" domanant trees, something like only
5%  (maybe a bit more if we consider the effects of random spacing of
the best trees) of the initial population need have really good growth
potential. When reading the growth of seedlings in a provenance trial,
a strain that produces a small number of superior trees might be
missed, but might be a good strain nevertheless.

   Dr. Charles Maynard at SUNY Syracuse has done some work with Norway
spruce, and he always emphasizes the importance of developing "land
races," that is a localized genetic strain produced by planting trees
from a good source, letting the trees grow to something like maturity,
and then collecting the seed of the remaining dominant trees to use in
subsequent plantings in the same or similar areas.  He has felt that
this may be more productive than doing trials of various European
sources, and then selecting the best based on a simple provenance
trial.

   The one limitation of this use of land races is sometimes a
specific stand one might want to use as a source of a land race, may
not have had the very basic potential of another.  For example, I
would much prefer to collect seed from the stand near Glady, WV than
the "Rothkugel." Anyway, with the growth of the "native trees only"
movement, interest in Norway spruce in this country has dropped off a
cliff!

   Yes, I digress.

   Anyway, I think I have pumped myself dry on this topic.  I am flat
out of ideas. But I would like to see othes with some insights to
chime in.  In the meantime, I will enjoy watching my pine groves grow,
and will be interested in finding good stands in which I--eventually
maybe--or others can measure outstanding trees. White pines are
wonderful even if they didn't grow to 250 feet.  But that idea is so
attractive, that I can understand why it is so hard for some of us to
abandon.

  --Gaines.
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