Pieter, 

 

I seemed to have dropped the thread.  I have downloaded a lot of the references 
you sent and will work through them as best I can.  It’s a very interesting 
list.  

 

But I haven’t responded to your list of plain spoken things to say about group 
selection.  So I will “lard” that below.  Lots going on here, health issues and 
what, but I want to staty with you on this if only for my own sanity.  

 

See below.

 

Nick

 

rom: Friam <[email protected]> On Behalf Of Pieter Steenekamp
Sent: Monday, August 11, 2025 7:06 AM
To: The Friday Morning Applied Complexity Coffee Group <[email protected]>
Subject: Re: [FRIAM] Group Selection IS a metaphor.

 

I think I’ve got a handle—maybe totally off the mark—on individual and group 
selection in human behavioural evolution. Here’s my take:

What is it? ChatNST==> Pieter: you come to me as an “expert” and I hope you 
understand that I am not (merely?) trolling when I say that the question is 
more interesting than the answer.  It reminds me of discussions about the hard 
problem of consciousness.  Its my “expert” opinion that, as in that case, the 
hard problem here is only hard for those people who want it to be hard.  For 
those who truly want to crack the problem, it is already answered, or its 
answer is extremely intricate and may only come after many years of balanced 
and careful research working out how both processes have left their mark. This 
case may be even harder than the consciousness case because it is an 
idiographic question, an attempt to explain a particularity, an individual 
historical event, human evolution.  I am not sure it is ever possible to 
“explain” a particularity, except insofar as one is unwilling to see it 
stripped of its uniqueness. So, then, how thoroughly are we committed to the 
idea of human uniqueness.  So that is what leads me to ask what hankering leads 
anyone to see the problem as hard.  I suspect that lying at the bottom of all 
of this is an ugly tangle of libertarian, liberal (in the classic sense) and 
progressive (including, perhaps some Marxism) thought which, if one is in that 
tangle, appears to make the question not only interesting but vital. Working 
through that tangle would be interesting but has little to do with the science. 
 Answering the scientific question is a long boring slog to work out the 
details of what we already know to be true.That IS my “expert” opinion and 
should be taken with a pound of salt. <==ChatNST
Human behaviour is a deeply complex phenomenon, shaped by two intertwined 
evolutionary processes: ChatNST==> Please try not to confound evolution with 
natural selection.  Any more than you would confound the theory OF anything 
with the thing it is a theory of.  Avoiding this confusion is not as easy as it 
sounds.  Do things fall because of gravity or is gravity the fact that things 
fall?  It can of course be either, but it can’t keep wobbling back and forth in 
the same discussion.  That is the logical flaw of equivocation. 

 

As usual, I speak with such “authority” and conviction because I know that 
others (Glen? DaveW? EricS? Russ?)  will protect you from me.  <== ChatNST

Biological evolution (based in our genes), where the unit of selection is the 
individual human.

Cultural evolution (based in memes), where the unit of selection is defined at 
the group level—though “group” can mean different things in different contexts.

 

ChatNST==> So, what do you take evolution to be apart from its most famous 
explainer, the differential replication of alternative heritable something?  
Can we really know which iron to choose without knowing where the green is? 
<==ChatNST



That’s it, in my simple words.

 

ChatNST==> Ah, if only these words were simple.  Thank you thank you for the 
report and bibliography.   <==ChatNST



Why do I say so?
I asked George to take a deep dive into the topic and produce a report, which 
you can view here: https://g.co/gemini/share/b046bfad5cc8 . I buy into its main 
argument—not claiming it’s “correct” in the absolute sense, but in my view it’s 
the best current explanation for the roots of human behaviour.

It draws on the latest research and, in the conclusion, puts it like this 
(paraphrased from the report):

The multilevel selection (MLS) framework, championed by David Sloan Wilson, 
integrates individual selection into a broader system of competing evolutionary 
forces within a nested hierarchy of genes, individuals, and groups. Extending 
this to cultural evolution, MLS offers a two-track model—genetic and 
cultural—that better explains human nature. By explicitly modelling the tension 
between within-group selfishness and between-group cooperation, it provides a 
richer explanation for complex social behaviours than alternatives like kin 
selection. The framework has implications beyond biology—touching religion, 
politics, and economics—and offers a unified, evolutionarily grounded lens on 
human sociality.

That’s why, for now, I’m hitching my wagon to this model.

 

References


Works cited


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2025,  
<https://eclass.uoa.gr/modules/document/file.php/PHS575/%CE%86%CF%81%CE%B8%CF%81%CE%B1/Okasha_Why%20won%27t%20group%20selection%20go%20away.pdf>
 
https://eclass.uoa.gr/modules/document/file.php/PHS575/%CE%86%CF%81%CE%B8%CF%81%CE%B1/Okasha_Why%20won%27t%20group%20selection%20go%20away.pdf

2.     Evolution of cooperation by multilevel selection - PNAS, accessed August 
11, 2025,  <https://www.pnas.org/doi/10.1073/pnas.0602530103> 
https://www.pnas.org/doi/10.1073/pnas.0602530103

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https://www.americanscientist.org/article/evolution-for-the-good-of-the-group

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11, 2025,  
<https://www.uvm.edu/~jfarley/EEseminar/readings/wilson-wilson.Rethinking20Sociobiology.inpress.pdf>
 
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https://www.cambridge.org/core/journals/behavioral-and-brain-sciences/article/reintroducing-group-selection-to-the-human-behavioral-sciences/634687DF831997525E21E5899B23CC8D

10.  Group selection - Wikipedia, accessed August 11, 2025,  
<https://en.wikipedia.org/wiki/Group_selection> 
https://en.wikipedia.org/wiki/Group_selection

11.  David Sloan Wilson's Group Selection Theory of Religion: Analysis and 
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<https://christianscholars.com/david-sloan-wilsons-group-selection-theory-of-religion-analysis-and-possible-christian-responses/>
 
https://christianscholars.com/david-sloan-wilsons-group-selection-theory-of-religion-analysis-and-possible-christian-responses/

12.  Eight Criticisms Not to Make About Group Selection - ResearchGate, 
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<https://www.researchgate.net/publication/51195160_Eight_Criticisms_Not_to_Make_About_Group_Selection>
 
https://www.researchgate.net/publication/51195160_Eight_Criticisms_Not_to_Make_About_Group_Selection

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- Journals, accessed August 11, 2025,  
<https://royalsocietypublishing.org/doi/10.1098/rspb.2024.3061> 
https://royalsocietypublishing.org/doi/10.1098/rspb.2024.3061

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- PMC, accessed August 11, 2025,  
<https://pmc.ncbi.nlm.nih.gov/articles/PMC11919500/> 
https://pmc.ncbi.nlm.nih.gov/articles/PMC11919500/

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11, 2025,  <https://jackbalkin.yale.edu/3-memetic-evolution> 
https://jackbalkin.yale.edu/3-memetic-evolution

16.  Memetics - Wikipedia, accessed August 11, 2025,  
<https://en.wikipedia.org/wiki/Memetics> https://en.wikipedia.org/wiki/Memetics

17.  Memetics - Principia Cybernetica Web, accessed August 11, 2025,  
<http://pespmc1.vub.ac.be/MEMES.html> http://pespmc1.vub.ac.be/MEMES.html

18.  pmc.ncbi.nlm.nih.gov <http://pmc.ncbi.nlm.nih.gov> , accessed August 11, 
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<https://pmc.ncbi.nlm.nih.gov/articles/PMC2607340/#:~:text=Genes%20and%20culture%20are%20two,acting%20back%20on%20the%20genome.>
 
https://pmc.ncbi.nlm.nih.gov/articles/PMC2607340/#:~:text=Genes%20and%20culture%20are%20two,acting%20back%20on%20the%20genome.

19.  Library: Susan Blackmore: Memetic Evolution - PBS, accessed August 11, 
2025,  <https://www.pbs.org/wgbh/evolution/library/07/2/l_072_05.html> 
https://www.pbs.org/wgbh/evolution/library/07/2/l_072_05.html

20.  The Meme Machine — by Susan Blackmore - Mimetic Theory, accessed August 
11, 2025,  <https://mimetictheory.com/the-meme-machine-by-susan-blackmore/> 
https://mimetictheory.com/the-meme-machine-by-susan-blackmore/

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<https://www.researchgate.net/publication/292001401_Memetics_A_dangerous_idea> 
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<https://en.wikipedia.org/wiki/Dual_inheritance_theory> 
https://en.wikipedia.org/wiki/Dual_inheritance_theory

23.  (PDF) Not By Genes Alone: How Culture Transformed Human ..., accessed 
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https://pmc.ncbi.nlm.nih.gov/articles/PMC3279745/

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<https://www.journals.uchicago.edu/doi/full/10.1093/bjps/axu047> 
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Research Portal, accessed August 11, 2025,  
<https://research-information.bris.ac.uk/files/34712898/Equivalence_MLS_and_IF_BJPS.pdf>
 
https://research-information.bris.ac.uk/files/34712898/Equivalence_MLS_and_IF_BJPS.pdf

27.  The Relation between Kin and Multilevel Selection: An Approach ..., 
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<https://www.journals.uchicago.edu/doi/10.1093/bjps/axu047> 
https://www.journals.uchicago.edu/doi/10.1093/bjps/axu047

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29.  Evolution of parochial altruism by multilevel selection - Max Planck 
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<http://web.evolbio.mpg.de/~garcia/preprints/07.pdf> 
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30.  The multilevel economic paradigm | INET Oxford, accessed August 11, 2025,  
<https://www.inet.ox.ac.uk/projects/the-multilevel-paradigm> 
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31.  Why Multilevel Selection Matters, accessed August 11, 2025,  
<https://scholarcommons.scu.edu/context/econ/article/1104/viewcontent/Field_Why_Multilevel_Selection_Matters.pdf>
 
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32.  Gene–culture coevolution and the nature of human sociality - PMC, accessed 
August 11, 2025,  <https://pmc.ncbi.nlm.nih.gov/articles/PMC3048999/> 
https://pmc.ncbi.nlm.nih.gov/articles/PMC3048999/

 

On Sun, 10 Aug 2025 at 09:41, Pieter Steenekamp <[email protected] 
<mailto:[email protected]> > wrote:

Nick (aka ChatNJT),

Are you still my George in this grand quest to truly “get” group selection?

A bit of backstory
When I get into a debate, my natural instinct is… well, to stay in my own head 
and not listen much at all. I first got called out on it nearly 50 years ago, 
back when I was a young engineer in management training. We did simulated 
negotiations, the instructor took notes, and the verdict was: “You didn’t hear 
a word they said.”

That kicked off my slow, sometimes painful, self-improvement journey. My 
instincts haven’t really changed (old habits die hard), but now I can force 
myself to switch gears — to really listen and understand the other person’s 
point before I jump in swinging.

I’ll admit, I lean toward believing in individual selection in human evolution. 
But I’m genuinely open to being convinced otherwise — and I want to really 
understand group selection before I form my final view.

I said I’d be spending some time learning with ChatGPT, and you kindly offered 
to be my “George” through ChatNJT. I’ve seen you active in this thread, but I’m 
not sure if our little backchannel on this is still alive or quietly drifting 
into the great beyond.

And hey, in the spirit of “why have a mind if we can’t change it?” — if you 
want to step back from this role, no hard feelings at all.

 

On Sat, 9 Aug 2025 at 06:49, Pieter Steenekamp <[email protected] 
<mailto:[email protected]> > wrote:

Nick, thanks for the document, I have downloaded it and will read it.

Next point, you do ask a lot of questions, Nick — and not the easy kind either. 
But fine, let’s dance.

"What is your hankering?"
I’m a simple creature. I just want to get a grip on what “group selection” 
really means for humans — simple enough to explain without a headache, but not 
so simple that it’s wrong. And, ideally, I’d like a reason to actually believe 
it exists.

"Where do you hope this will all come out?"
Same answer, really. I trust my brain enough to think I can untangle 
complicated stuff… eventually. My hope is just to reach that magical “ohhh, 
that’s what it means” moment.

"What would group selection look like in human beings?"
Now you’re hitting the nerve. I can’t answer that — which is exactly why I’m 
here poking at the question.
Right now, it feels at odds with the simple elegance of evolution, which (as 
ChatGPT put it) goes like this:

Evolution is the gradual change of replicators — things that make copies of 
themselves — over time. Sometimes the replicator exists inside a temporary form 
(like an organism, idea, or machine) that competes with others. Variations that 
help it succeed in making more copies become more common, shaping the system 
over time.

And here’s my snag: I see humans as one big messy group, not a bunch of smaller 
competing groups. So where’s the competition? Clearly I’m missing a big chunk 
of the story — and I want to find it.

"Would you approve or disapprove?"
I’m not here to pass moral verdicts. I just want to figure it out before 
deciding whether to even have an opinion.

"What is a group? Is a species a group? Is a race a group? Is a village a 
group?"
And there’s the heart of my confusion. Right now, my brain says: “Well, all 
humans are one group, right?” — which doesn’t fit neatly with my current 
picture of evolution. So the plan is simple: swap ignorance for understanding, 
and hopefully keep the coffee hot while I do it.

 

On Fri, 8 Aug 2025 at 23:52, <[email protected] 
<mailto:[email protected]> > wrote:

Great Peiter, 

 

But you didnpt answer my question.  I know it’s the hardest kind of question to 
answer, but give it a go.  What is your hankering?  Where do you hope this will 
all come out?   What would group selection look like in human beings?  Would 
you approve of it or disapprove of it?  What is a group, after all?  Is a 
species a group?  Is a race a group? Is a village a group? Etc.  

 

DS Wilson I think lost interest in the question that most interested me (what 
are the elemental forces that led to the evolution of complex organisms) and 
became more interested in in the forces that lead to human groupish behavior.  
To me human groupishness seems wildly overdetermined.  Its like asking why is 
the pope a Christian.  But that’s a wildly unsatisfying answer to some one who 
is genuinely surprized to find that the pope is indeed a Christian.  

 

Lets go back and forth like this for a few more exchanges.

 

Meantime, I enclose a short article in BBS that reprises a much larger article 
by W and S.   I have a pdf of the larger article on my hard drive and will send 
it to you when I figure out how to bypass friam’s restrictions on large files. 

 

But please don’t let that get in the way of you taking a shot at answers to the 
questions I posed.

 

Nick

.  

 

From: Friam <[email protected] <mailto:[email protected]> > On 
Behalf Of Pieter Steenekamp
Sent: Friday, August 8, 2025 4:21 PM
To: The Friday Morning Applied Complexity Coffee Group <[email protected] 
<mailto:[email protected]> >
Subject: Re: [FRIAM] Group Selection IS a metaphor.

 

Nick,

Too good to miss — I’m in. Lead me into the jungle of group selection, 
especially the human variety.

What I’m after: a clear, simple (but not dumbed-down) take on what group 
selection in humans is, and why it might explain our behaviour better than 
individual selection alone.

Happy to start at the very beginning — dawn of the argument, cave paintings, 
whatever you think works.

And yes, send me that Famous Great Amateur reading list. I promise to read it 
with respect… and just enough suspicion to keep it fun.

 

On Fri, 8 Aug 2025 at 17:05, Nicholas Thompson <[email protected] 
<mailto:[email protected]> > wrote:

Hi, Pieter, 

 

Let me be a George to you as you explore this topic.  I will try to respond off 
hand, quickly, and unself-consciously as you think along.  I think this whole 
topic is fascinating both substantively, and historically.   The literature 
seems to track (or lead?) the Zeitgeist so precisely from post war peace-nikery 
(Wynne-Edwards), to the revanchist academic Reaganism (Williams-Dawkins), to 
chaos (evodevo). It's really hard to take the whole argument seriously once one 
begins to understand how complex and multi layered are the mechanisms by which 
parents do and dont resemble their children.   One of the tools to thinking 
straight is to own up to one's hankerings before one dives into the literature. 
 What are you hoping to find?  Post war peace-nikery was covertly deistic,  
hoping to find that there was some sort of over arching regulatory agency that 
would keep the species and the planet safe.  Academic Reaganism said good luck 
with that!   Success is virtue.  And then evodevo, the bull in the china shop 
of that whole argument.  I recommend reading the biologist, Sean B. Carroll, 
(not the physicist), Endless forms most beautiful, and The making of the 
fittest.   It's really hard to take the whole argument seriously once one 
begins to understand how complex and multi layered are the mechanisms by which 
parents do and dont resemble their children. That there is any resemblance at 
all begins to seem like some sort of miracle.  Or perhaps just momentum.  One 
hankering that misleads us is naturalism, the idea that we can find some sort 
of MORAL guidance in the way things are.  Is the opposite hankering, 
existentialism?  The belief that what makes humans special is their power to 
CHOOSE.  You should remember that I am not a philosopher and am, in fact, an 
amateur in all things.  

 

"Any time you want to explore this issue, I  am here ready to help.  Would you 
like suggestions of articles to read by that Famous Amateur, Nick Thompson? "

 

signed, 

 

ChatNST

 

 

 

On Fri, Aug 8, 2025 at 5:19 AM Pieter Steenekamp <[email protected] 
<mailto:[email protected]> > wrote:

Thanks, Nick. Just like you struggled to get your head around entropy, I’m 
battling to wrap my mind around how the basic but very powerful mechanism of 
evolution works in human groups. I can easily understand individual human 
selection, or even group selection in swarming insects where only the queen has 
babies.

I think I’ll take a page from your book and work with George to help guide me 
through this learning journey. Every now and then, I might check in with you 
and others here for a chat or to ask a question.

The only catch is that I’ve just started a really exciting AI project, so I 
might not have much time for my group-level evolution journey — but I’ll try to 
keep it going.

 

On Fri, 8 Aug 2025 at 03:40, <[email protected] 
<mailto:[email protected]> > wrote:

Thanks Pieter, 

 

Sorry I have taken so long to get back to you.  If FRIAM ever started a 
journal, it should be called “the emperors new clothes”.  We are not committed 
to anything if not to the validity of an “amateur’s” perspective.  As people 
will be quick to tell you, mine has always been of that sort. 

 

If I read you carefully, the position you take is that laid out in Dawkins The 
Extended Phenotype – that the genes are the basic unit of selection.  But as 
Dave Wilson has been pointing out for years, Who made that decision?   For one 
thing, as epigenic studies have made clear, when one looks in detail, it is 
really hard to find a thing that is exactly the gene.  For another, that 
decision runs the risk of confusing the the thing that is selected with the 
forces that are selecting it.  Whatever level you care to calculate the impact 
of selection, it is differential group success that is driving selection or it 
is not group selection.  And if it  is differential group success that is 
driving selection, then it is group selection.  I think you might quite enjoy 
The Extended Phenotype.   For a whild ride, have a look at Elliott Sober and D. 
S. Wilson’s Reintroducing Group Selection to the human behavioral sciences.  
There is a wonderful metaphor in there about two riders riding three horses.  
It was the article that broke the tide for me.  I had been totally up Dawkins 
ass for the preceding 20 years. 

 

Here is the citation, courtesy og George Patrick Tremblay IV  

 

Wilson, D. S., & Sober, E. (1994). Reintroducing group selection to the human 
behavioral sciences. Behavioral and Brain Sciences, 17(4), 585–608. 
https://doi.org/10.1017/S0140525X00036104 en.wikipedia.org+15philpapers.org+15 
<https://philpapers.org/rec/WILRGS?utm_source=chatgpt.com> ….

 

Nick 

 

From: Friam <[email protected] <mailto:[email protected]> > On 
Behalf Of Pieter Steenekamp
Sent: Wednesday, August 6, 2025 12:55 AM
To: The Friday Morning Applied Complexity Coffee Group <[email protected] 
<mailto:[email protected]> >
Subject: Re: [FRIAM] Group Selection IS a metaphor.

 

Nick, I'm genuinely impressed. Honestly, I feel a bit out of my depth trying to 
respond meaningfully on this topic.

So please take my reply in the same spirit I’d expect a response from my 
10-year-old grandchild when debating computer programming with me. The gap 
between your understanding of evolution and mine feels about that wide.

That said, I’d still like to offer a response to your group selection 
argument—fully aware that it may come across as amateurish, and I'm okay with 
that.

Here's the question I’m grappling with:

Is the following valid?
Genes as the Unit of Selection:
Modern evolutionary theory generally views genes as the primary unit of 
selection. Natural selection acts on individuals, and the success of an 
individual is ultimately determined by the genes they carry. 
Group Selection as a Modifier:
Group selection can be seen as a process that influences the expression of 
genes. For example, if a group-level trait (like cooperative behavior) is 
advantageous, then genes that promote that behavior will be favored, even if 
those genes also have individual-level costs.

 

On Wed, 6 Aug 2025 at 00:12, Prof David West <[email protected] 
<mailto:[email protected]> > wrote:

Nick,

 

I wish to embody the fear of being dragged away from what you think you are 
supposed to be doing, to be engaged in the topic you raise in your paper.

 

I have read the paper before and, as then, I find it meritorious, well written, 
and reasonable in argument. I am, basically, convinced.

 

However; two points:

 

First, your use of the concept, "metaphor," is the way that I use the term, in 
a manner that glen pointed out is inconsistent with the literal definition of 
the term. I speak of metaphor when there is some thing of which I think I know 
something and I have a suspicion that some other thing might be of the same 
ilk. I use what I think I know to craft a 'model', one that suggests particular 
points and particular relations that, if my suspicion is correct, will have 
direct analogs in the unknown thing. I check them out individually and in 
combinations and, if substantiated, confirm my suspicion. If unconfirmed, the 
metaphor is refuted.

 

This seems to me to be what you are doing in the paper, albeit it more 
abstractly and academically. Please correct me if wrong.

 

Second, and here is the real time sink, would it be possible to make your ideas 
concrete, real groups with actual history and demonstrated differential 
"success." If you were amenable to such a conversation, I would propose the 
Mormons as a test case.

 

One of 20 or so "religions"/"societies" to emerge from the "Burnt Over 
District" of western New York. The only one still extant.

 

Disproportionately successful, (in material and social terms), to their 
neighbors. Smith was living in a two-story New England style home while down 
the road, Abe Lincoln, was living in a log cabin with mud floor.

 

A schism immediately after Smith's death, with the Reformed LDS barely evident 
while the main group flourished. (Last time I checked, Mormonism and Sokka 
Gokai, in Japan, were the two fastest growing religions.)

 

In Utah there was a concerted effort to spawn multiple small groups by sending 
out colonies. Because each group was originally "seeded" with four or five 
families, you get a strong genetic/heritance component as well as "traits." (It 
is still possible to identify what part of Utah someone is from (especially 
females) by their physical appearance.)

 

Some interesting "adaptations" at the trait level, e.g., when Smith was alive 
blacks were included in the community and held the priesthood—something that 
Missourians, at the time, could not abide. Brigham Young 'suspended' (restored 
in 1978 with the admission that the suspension was not for theological, but 
merely political reasons) black priesthood membership and gave up polygamy (de 
jure only) to appease the Federal Government and avoid a second martyrdom.

 

davew

 

 

On Tue, Aug 5, 2025, at 1:10 PM, Nicholas Thompson wrote:

Dear Colleagues in FRIAM,

Sometimes, if I am going to get anything done, I just have to ignore Friam, and 
keep my head down, and work at the thing I am working at.  It always seems, on 
that occasion, that you-guys dangle in front of me some enticing topic so I 
must scream and put my fingers in my ears to keep focus on my work.  So it was 
that when I decided I must fish or cut bait on entropy or it would take me to 
my grave, that almost immediately you-guys started not one but two 
conversations close to my heart: on the centrality of metaphor to science and 
on the group selection controversy. 

A couple of decades ago I brought those two interests together in  a paper 
called “Shifting the Natural Selection  Metaphor to the Group Level.  There are 
two things about this paper that make it salient for me.  The first is that I 
think it is the best paper I ever wrote.  The second is that for each of the 
two people whom I most hoped to reach when I wrote it, D. S. Wilson and Elliott 
 Sober, it is a piece of  crap. In it, I try to show that the problem with 
metaphors is not with their use in scientific thinking: on the contrary, it is 
with their ill-disciplined use.  Metaphors need to be worked in a systematic 
way, not simply flung out in a gust of poetic exuberance.  This lesson  I try 
to teach by working the natural selection metaphor in a systematic way to show 
that if it had been treated seriously in the first place, the whole dispute 
about group selection might have been  avoided.  Thus the paper is not only 
arrogant, but meta-arrogant.  

Nothing is more pitiable than the retired academic who would do anything to 
have anybody read his moribund essays.  But, alas, I simply am such a person.  
So, I am attaching a copy of the paper  in the hope that it will have some 
value to you within the context of your two discussions. 

Mumble,

Nick

 

 

--

Nicholas S. Thompson

Emeritus Professor of Psychology and Ethology

Clark University

[email protected] <mailto:[email protected]> 

https://wordpress.clarku.edu/nthompson

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Attachments:

*       Shifting the natural selection metaphor to the group level.pdf
*       Shifting the natural selection metaphor to the group level.pdf

 

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-- 

Nicholas S. Thompson

Emeritus Professor of Psychology and Ethology

Clark University

[email protected] <mailto:[email protected]> 

https://wordpress.clarku.edu/nthompson

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