Thanks a lot, Rawat ji

---------- Forwarded message ---------
From: GOPAL SINGH Rawat <[email protected]>
Date: Tue, 12 Apr 2022 at 17:53
Subject: Re: What is Saxifraga diversifolia ?
To: J.M. Garg <[email protected]>


Dear Garg ji,

Very pertinent question. Let's see what others have to say.

As indicated by Eflora of China and also Vol 1, Flora of Nepal, in S.
parnassifolia the cauline leaves are more or less amplexicaule and the
basal leaves resemble that of Parnassia. In S. diversifolia the cauline
leaves ovate and not amplexicaule.

Earlier illustrations and photos could be mixed and include S.
parnassifolia which was considered a variety of S. diversifolia for a long
time.

Having said this, I need to see it more closely in the field.

Regards,

G S Rawat




On Tue, Apr 12, 2022 at 2:48 PM J.M. Garg <[email protected]> wrote:

> Dear members,
> After going through images at Saxifraga parnassifolia
> <https://efloraofindia.com/2011/12/13/saxifraga-parnassifolia/>, I got
> confused after seeing the variations in images herein.
> On searching in detail, I could not find the real difference between the
> two and inf. being contradictory at various places.
> Flora of China
> <http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=305208&key_no=6>
> gives the following keys:
> 24 (23)
> <http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=305208&key_no=6#KEY-6-23>
>  At
> least some cauline leaves ovate, base cordate, ± amplexicaul; sepals erect
> to spreading.   (25)
> <http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=305208&key_no=6#KEY-6-25>
> + Cauline leaves narrowly elliptic, base rounded, not amplexicaul; sepals
> reflexed.   (26)
> <http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=305208&key_no=6#KEY-6-26>
>
> 25 (24)
> <http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=305208&key_no=6#KEY-6-24>
>  Petals
> to 8 mm   29 S. parnassiifolia
> <http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=241000345>
> + Petals at least 8 mm   28 S. sphaeradena
> <http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=200010396>
>
> 26 (24)
> <http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=305208&key_no=6#KEY-6-24>
>  Inflorescence
> cymose, 2- or 3-flowered, or flower solitary; basal leaf blade 1.1-2 cm
> 23 S. dianxibeiensis
> <http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=200010251>
> + Inflorescence corymbiform, 5-17-flowered; basal leaf blade 1.5-5 cm
> 22 S. diversifolia
> <http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=200010256>
>
> However, it looks contradictory in Plant illustrations
> <http://www.plantillustrations.org/species.php?id_species=912173> one
> <http://www.plantillustrations.org/illustration.php?id_illustration=80744&id_taxon=8103&mobile=0&SID=pqmo7ph77jo3gk5g0id89087d5&language=English&thumbnails_selectable=0&selected_thumbnail=0&query_type=species&query_broad_or_restricted=broad&group=0&lay_out=0&uhd=0>
> and two
> <http://www.plantillustrations.org/illustration.php?id_illustration=4128&id_taxon=8103&mobile=0&SID=pqmo7ph77jo3gk5g0id89087d5&language=English&thumbnails_selectable=0&selected_thumbnail=0&query_type=species&query_broad_or_restricted=broad&group=0&lay_out=0&uhd=0>
>  of Saxifraga
> diversifolia.
>
> Here is the description of Saxifraga diversifolia from IBIS Flora from
> Flora of British India
> <https://flora.indianbiodiversity.org/content/saxifraga-diversifolia-wall-ex-ser-0>
>  (could
> not find it for Saxifraga parnassifolia):
>
> *Wall. in Sternb. Saxifrag. Suppl, t.* 22 ; flowers corymbose rarely 3-2,
> sepals ovate-oblong small, cauline leaves sessile often cordate and
> amplexicaul,* radical leaves long-petioled often large.* Wall. Cat. 452 ;
> DC. Prodr. iv. 44 ; Hook. Lond. Journ. Bot. iv. t. 21 ; H. f. & T. in
> Journ. Linn. Soc. ii. 70 ; Engler Monog. Saxifrag. 125. S. parnassifolia,
> Wall. Cat. 451, partly ; Don in Trans. Linn. Soc. xiii. 405 ; DC. Prodr.
> iv. 25 ; Sternb. Saxifrag. Suppl, t. 25. S. Moorcroftiana, Wall. Cat. 453 ;
> Sternb. Saxifrag. Suppl, t. 24. S. lysimachoides, Klotzsch in. Reis. Pr.
> Waldem. Bot. t. 42.
> Alpine Himalaya ; from KASHMIR to BHOTAN, alt. 9000-17,000 ft., abundant.
> Stems 8-18 in. high, leafy, glandular, often finally glabrous. *Radical
> leaves long-petioled, narrowed into the petiole *or (var. parnassifolia)
> ovate, subcordate ; cauline 1/2-2 in., ovate or broad oblong. Sepals at
> length reflexed. Petals with 4 stalked glands at their base, 1.1/2—2 times
> the sepals. Styles very short. Capsule ovate-oblong. Seeds angular, testa
> smooth, somewhat plaited.
>
> Here is what Flora of China says:
> Saxifraga diversifolia
> <http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=200010256>:
> Herbs perennial, 16-43 cm tall. Stem proximally brown crisped villous or
> glabrous, distally shortly glandular hairy. *Basal leaves with petiole
> 3-9 cm, abaxially and marginally brown villous; leaf blade ovate-cordate to
> narrowly ovate, 1.5-5 × 1.2-2.6 cm, abaxially and marginally brown
> pubescent, **adaxially** glabrous or brown pilose near base, base
> cordate, apex acute.* Cauline leaves 8-12; petiole 0.5-4 cm, brown
> crisped villous; leaf blade subcordate or ovate-cordate to narrowly ovate,
> 1-6.3 × 0.4-4 cm, base cordate or subcordate, apex obtuse or acute;
> proximal leaf blades larger, abaxially and marginally brown pilose or
> glandular pilose, adaxially glabrous or subglabrous; distal leaf blades
> smaller, usually glabrous on both surfaces, glandular hairy at margin. Cyme
> corymbose, 3-14 cm, 5-17-flowered; pedicels 6-12 mm, glandular hairy.*
> Sepals reflexed, *narrowly to broadly ovate, 3-4.2 × 1.3-3.5 mm,
> abaxially glandular hairy or glabrous on both surfaces, veins 3(-5), not or
> partly confluent at apex, margin membranous, glandular ciliate, apex obtuse
> or acute, rarely erose. Petals yellow, elliptic, obovate, or ovate to
> narrowly so, rarely oblong, 5-8 × 2-5 mm, usually not callose, rarely
> 4-6-callose, (3-)5-7(-9)-veined, base narrowed into a claw 0.5-1.3 mm, apex
> obtuse. Stamens 4-5.6 mm. Ovary subsuperior, ovoid, 3-4.2 mm; styles 1-1.6
> mm. Fl. and fr. Aug-Oct. 2n = 16, (?20).
>
> Forests, forest margins, scrub, alpine meadows, rock crevices; 2700-4300
> m. W Sichuan, SE Xizang, N Yunnan [Bhutan, Kashmir, Myanmar, Nepal, Sikkim].
>
> *Saxifraga diversifolia* is extremely polymorphic. Many of the variants
> have been published as species, which are accepted as such in the present
> account because a very narrow species concept has been adopted in
> classifying the variation. However, many botanists would prefer to see
> rather more "lumping," and comments relating to how this might be done are
> made in relevant places.
>
>
> *Saxifraga parnassiifolia
> <http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=241000345>* D.
> Don <http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=241000345>
> :
> *Saxifraga diversifolia* Wallich ex Seringe var. *parnassiifolia* (D.
> Don) Seringe;
> Herbs perennial, cespitose, 11.5-24 cm tall. Stem branched, densely brown
> crisped glandular villous. *Basal leaves with petiole 1.3-2.7 cm, crisped
> glandular villous; leaf blade cordate-ovate, 1.5-4 × 1.3-3 cm, crisped
> villous on both surfaces and at margin or glabrous adaxially, base cordate,
> apex acute.* Cauline leaves 6 or 7, sessile, ovate to cordate, 0.9-3.5 ×
> 0.5-2.3 cm, base cordate, semiamplexicaul, apex obtuse or acute; proximal
> leaves glandular villous on both surfaces and at margin or glabrous
> adaxially; distal leaves shortly glandular hairy on both surfaces, margin
> shortly glandular hairy and glandular villous. Pleiochasium 2-2.5 cm,
> 6-11-flowered; pedicels shortly glandular hairy. *Sepals erect to
> spreading, *ovate to broadly so, 3-4 × 1.5-2.9 mm, shortly brown
> glandular hairy abaxially, adaxially distally, and at margin and apex,
> veins 5-7, confluent into a verruca at apex. Petals yellow, obovate to
> broadly ovate, 5-7.9 × 3.5-4 mm, subcolumnar 4-6-callose, not or
> indistinctly 2- or 3-callose, 5-7-veined, base contracted into a claw
> 0.6-1.7 mm, apex obtuse. Stamens 2.5-4 mm. Ovary subsuperior, ovoid,
> 1.6-2.5 mm; styles 1-2 mm. Fl. Jul-Sep. 2n = 16.
>
> *Abies* forest margins, slopes; 2700-4000 m. S Xizang (Nyalam Xian, Zayü
> Xian) [Bhutan, India, Nepal, Sikkim].
>
> *Saxifraga parnassiifolia* var. *obscuricallosa*, which was accepted in
> FRPS, cannot be distinguished reliably because of intermediates.
>
> *Can anyone solve this mystery or give clear cut and verifiable keys
> between the two?*
> --
> With regards,
> J.M.Garg
>


-- 
With regards,
J.M.Garg

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