On Thu, 14 Oct 2021, Rich Shepard wrote:

Looks like the issue is with Keefer, et al. 2008.

Here's the biblatex entry for that paper:
@Article{Keefer2008,
  author   = {Keefer, M.L. and Boggs, C.T. and Peery, C.A. and Caudill, C.C.},
  journal  = {North American Journal of Fisheries Management},
  title    = {Overwintering Distribution, Behavior, and Survival of Adult 
Summer Steelhead: Variability among Columbia River Populations},
  year     = {2008},
  pages    = {81--96},
  volume   = {28},
  abstract = {Unlike most anadromous salmonids, summer steelhead Oncorhynchus
                  mykiss overwinter in rivers rather than the ocean for 6--10
                  months prior to spring spawning. Overwintering in rivers may
                  make summer steelhead more vulnerable to harvest and other
                  mortality sources than are other anadromous populations, but
                  there has been little systematic study of this life history
                  strategy. Here, we used a large-scale radiotelemetry study
                  to examine the overwintering behaviors and distributions of
                  26 summer steelhead stocks within the regulated lower
                  Columbia-Snake River hydrosystem. Over 6 years, we monitored
                  5,939 fish, of which 3,399 successfully reached spawning
                  tributaries or the upper Columbia River basin and were
                  assigned to specific populations. An estimated 12.\% of fish
                  that reached spawning areas overwintered at least partially
                  within the hydrosystem (annual estimates = 6.8--19.6\%),
                  while the remainder overwintered in tributaries. Across all
                  populations, later-arriving fish were more likely to
                  overwinter in the hydrosystem; overwintering percentages
                  ranged from less than 1\% for fish tagged in June to over
                  40\% for those tagged in October. Proportionately more
                  interior-basin steelhead (Clearwater, Salmon, and Snake
                  River metapopulations) overwintered in the hydrosystem than
                  did fish from lower-river populations. Steelhead were
                  distributed in mixed-stock assemblages throughout the
                  hydrosystem during winter, usually in reservoirs closest to
                  their home rivers but also in nonnatal tributaries.
                  Overwintering fish moved upstream and downstream between
                  reaches in all months; a nadir occurred in early January and
                  peak egress into spawning tributaries was in March. The
                  estimated survival to tributaries was higher for fish that
                  overwintered in the hydrosystem (82\%) than for fish that
                  did not (62\%); this difference was largely attributable to
                  low winter harvest rates. Our results suggest that large
                  main-stem habitats, including reservoirs, may be widely used
                  by overwintering summer steelhead. The complex migration
                  behaviors of steelhead indicate both the potential for
                  adaptation and possible susceptibility to future river
                  environment changes.},
  keywords = {fish, salmon, steelhead, trout, rivers, Columbia River, 
distribution, behavior, survival, adults},
}

Are the '%' symbols not correctly escaped?

Rich
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