Does your SDS-PAGE loading buffer contain a reducing agent like beta
mercaptoethanol? That could be responsible for the difference between
your SDS-PAGE and HPLC results.


ho

On Fri, Mar 26, 2010 at 5:01 PM, CCP4BB automatic digest system
<[email protected]> wrote:
> There are 5 messages totaling 490 lines in this issue.
>
> Topics of the day:
>
>  1. TCEP effect on protein (4)
>  2. 3 PhD studentships available immediately
>
> ----------------------------------------------------------------------
>
> Date:    Thu, 25 Mar 2010 22:51:30 -0800
> From:    megha goyal <[email protected]>
> Subject: TCEP effect on protein
>
> Hi all,
>
> My protein is relatively pure except the dimer. i used 10 mM TCEP to reduce
> it (directly added 1 M TCEP to make final volume of 10mM and kept at room
> temperature for 10 mins] then i dialysed the protein sample to remove TCEP
> [dialysis buffer is Na acetate pH 4.0].
>
> On performing SDS PAGE analysis after dialysis of the protein we are getting
> no dimer band, only band of our protein is observed and same is the case
> with UV reading there is no change in it. But the HPLC analysis of the
> protein shows two peaks instead of one peak as observed before TCEP
> treatment. what can be the reason for this. Kindly guide. i need the protein
> to formulate and conduct stability studies on the sample. the protein we
> obtain after IEX is pure except the dimer and i do not want to go for SEC as
> it greatly reduces protein content and also is quite time consuming. any
> light on what is happening will bevery useful.
>
> thanks and regards
>
> ------------------------------
>
> Date:    Fri, 26 Mar 2010 08:37:43 +0100
> From:    Ganesh Natrajan <[email protected]>
> Subject: Re: TCEP effect on protein
>
>
>
> Hi Megha,
>
> The two peaks on the HPLC indicate that your protein is
> existing in a monomer-dimer equilibrium in solution. The dimerisation is
> most probably caused by disulphide bridges. The use of TCEP is breaking
> those disulphides and that is causing the equilibrium to move towards the
> monomeric state. However, when the TCEP is dialysed out, the disulphides
> start forming again and this is causing the equilibrum to move towards the
> dimeric state, a process clearly hastened by the strongly oxidising pH 4 of
> the dialysis buffer.
>
> Now it all depends on what you want to do. If you
> want to use the protein in a (largely) monomeric form, I would recommend
> that you don't dialyse out the TCEP.
>
> regards
>
> Ganesh
>
>
> **********************************************
> Blow, blow, thou winter
> wind
> Thou art not so unkind
> As man's ingratitude;
> Thy tooth is not so
> keen,
> Because thou art not seen,
> Although thy breath be rude.
>
> -William
> Shakespeare
> **********************************************
>
> On Thu, 25 Mar
> 2010 22:51:30 -0800, megha goyal  wrote:  Hi all,   My protein is
> relatively pure except the dimer. i used 10 mM TCEP to reduce it (directly
> added 1 M TCEP to make final volume of 10mM and kept at room temperature
> for 10 mins] then i dialysed the protein sample to remove TCEP [dialysis
> buffer is Na acetate pH 4.0].   On performing SDS PAGE analysis after
> dialysis of the protein we are getting no dimer band, only band of our
> protein is observed and same is the case with UV reading there is no change
> in it. But the HPLC analysis of the protein shows two peaks instead of one
> peak as observed before TCEP treatment. what can be the reason for this.
> Kindly guide. i need the protein to formulate and conduct stability studies
> on the sample. the protein we obtain after IEX is pure except the dimer and
> i do not want to go for SEC as it greatly reduces protein content and also
> is quite time consuming. any light on what is happening will bevery useful.
>  thanks and regards
>
> --
>
> ------------------------------
>
> Date:    Fri, 26 Mar 2010 08:49:55 +0100
> From:    Matthias Zebisch <[email protected]>
> Subject: Re: TCEP effect on protein
>
> Hi Ganesh and Mega!
>
> I do not agree with Ganesh. I assume, Megha, that truly reversed
> phaseHPLC was used. This is a denaturing method and the natural
> disulfide should not form again during the run.
> Also pH 4 can not be described "oxidizing". Actually, reduced proteins
> are often dialyzed against acidic buffers to prevent disulfide formation
> via the thiolate anion.
> Still, a reducing agent may be used during the run?
>
> Sorry that I can not offer a solution to the real problem. More
> experimental details may be necessary.
>
> Bets regards, Matthias
>
> Am 3/26/2010 8:37 AM, schrieb Ganesh Natrajan:
>>
>> Hi Megha,
>>
>> The two peaks on the HPLC indicate that your protein is existing in a
>> monomer-dimer equilibrium in solution. The dimerisation is most
>> probably caused by disulphide bridges. The use of TCEP is breaking
>> those disulphides and that is causing the equilibrium to move towards
>> the monomeric state. However, when the TCEP is dialysed out, the
>> disulphides start forming again and this is causing the equilibrum to
>> move towards the dimeric state, a process clearly hastened by the
>> strongly oxidising pH 4 of the dialysis buffer.
>>
>> Now it all depends on what you want to do. If you want to use the
>> protein in a (largely) monomeric form, I would recommend that you
>> don't dialyse out the TCEP.
>>
>> regards
>>
>> Ganesh
>>
>> **********************************************
>> Blow, blow, thou winter wind
>> Thou art not so unkind
>> As man's ingratitude;
>> Thy tooth is not so keen,
>> Because thou art not seen,
>> Although thy breath be rude.
>>
>> -William Shakespeare
>> **********************************************
>>
>> On Thu, 25 Mar 2010 22:51:30 -0800, megha goyal <[email protected]>
>> wrote:
>>
>>     Hi all,
>>     My protein is relatively pure except the dimer. i used 10 mM TCEP
>>     to reduce it (directly added 1 M TCEP to make final volume of 10mM
>>     and kept at room temperature for 10 mins] then i dialysed the
>>     protein sample to remove TCEP [dialysis buffer is Na acetate pH 4.0].
>>     On performing SDS PAGE analysis after dialysis of the protein we
>>     are getting no dimer band, only band of our protein is observed
>>     and same is the case with UV reading there is no change in it. But
>>     the HPLC analysis of the protein shows two peaks instead of one
>>     peak as observed before TCEP treatment. what can be the reason for
>>     this. Kindly guide. i need the protein to formulate and conduct
>>     stability studies on the sample. the protein we obtain after IEX
>>     is pure except the dimer and i do not want to go for SEC as it
>>     greatly reduces protein content and also is quite time consuming.
>>     any light on what is happening will bevery useful.
>>     thanks and regards
>>
>> --
>>
>
>
> --
> ****************************************************
> Dr. Matthias Zebisch
> Universität Leipzig
> Biotechnologisch-Biomedizinisches Zentrum
> Strukturanalytik von Biopolymeren
> Deutscher Platz 5
> 04103 Leipzig
> Germany
> Phone: 0049-341-97-31323 (lab) -31312 (office)
> Fax  : 0049-341-97-31319
> email: [email protected]
> ****************************************************
>
> ------------------------------
>
> Date:    Fri, 26 Mar 2010 10:53:07 +0000
> From:    "Hough, Mike" <[email protected]>
> Subject: 3 PhD studentships available immediately
>
> Dear CCP4bb,
>
> The following three studentship positions are available immediately at the 
> University of Liverpool. Could you please pass these details on to any 
> students in your departments or elsewhere who may be interested? Replies to 
> the email addresses listed below, rather than to me please.
>
> Thanks,
>
> Mike
>
>
> School of Biological Sciences, University of Liverpool
>
> 4-year joint BBSRC PhD Studentships – applications invited immediately
>
> Three PhD studentships are available in the Molecular Biophysics Group for 
> research in the area of x-ray structural biology using the most advanced 
> synchrotron radiation sources and the new light sources in the form of x-ray 
> free electron lasers (XFELs). The studentships involve collaborations with 
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> Japan. The successful candidates will have the exciting opportunity of 
> spending up to 2 years at these premier science facilities working at the 
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> project descriptions are given below:
>
> “Long wavelength X-ray diffraction experiments for structural studies from 
> biological redox systems”
> [Strange, UoL & Wagner, DIAMOND]
> see: http://www.biophysics.liv.ac.uk/PhD_DLS.pdf
> contact [email protected]
>
> “Metalloproteomics: Structure and function studies of metalloproteins of 
> biomedical importance”
> [Hasnain, UoL & Shiro, RIKEN]
> see: http://www.biophysics.liv.ac.uk/PhD_RIKEN_Metalloprotein.pdf
> contact: [email protected]
>
> “New Science exploration from XFEL: A new paradigm for structural 
> visualisation of macromolecules”
> [Grossmann, UoL & Ishikawa, RIKEN]
> see: http://www.biophysics.liv.ac.uk/PhD_RIKEN_XFEL.pdf
> contact: [email protected]
>
> ------------------------------
>
> Date:    Fri, 26 Mar 2010 09:40:48 -0700
> From:    James Holton <[email protected]>
> Subject: Re: TCEP effect on protein
>
> Yes, it is possible that the disulfide is just re-forming when the TCEP
> is gone.  pH 4 is not favorable for the oxidation, but does not prohibit
> it either.  Especially if there are traces of metal ions around.  I
> learned this the hard way, as trace metals can catalyze the oxidation of
> methionine and selenomethionine as well.  In my hands, adding a pinch of
> EDTA (~1 mM final conc.) to the fraction coming off the HPLC stabilized
> the reduced species.  Since the HPLC I was using at the time was made of
> metal (and showed signs of rust around some of the fittings), it is
> perhaps not surprising that I had a few ppm of Fe++ in the mobile phase.
>
> -James Holton
> MAD Scientist
>
> megha goyal wrote:
>> Hi all,
>>
>> My protein is relatively pure except the dimer. i used 10 mM TCEP to
>> reduce it (directly added 1 M TCEP to make final volume of 10mM and
>> kept at room temperature for 10 mins] then i dialysed the protein
>> sample to remove TCEP [dialysis buffer is Na acetate pH 4.0].
>>
>> On performing SDS PAGE analysis after dialysis of the protein we are
>> getting no dimer band, only band of our protein is observed and same
>> is the case with UV reading there is no change in it. But the HPLC
>> analysis of the protein shows two peaks instead of one peak as
>> observed before TCEP treatment. what can be the reason for this.
>> Kindly guide. i need the protein to formulate and conduct stability
>> studies on the sample. the protein we obtain after IEX is pure except
>> the dimer and i do not want to go for SEC as it greatly reduces
>> protein content and also is quite time consuming. any light on what is
>> happening will bevery useful.
>>
>> thanks and regards
>
> ------------------------------
>
> End of CCP4BB Digest - 25 Mar 2010 to 26 Mar 2010 (#2010-82)
> ************************************************************
>

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