People,

I a sorry my suggestions last night were not better articulated.  They were 
posted at 3 AM while I was up from not feeling well. There really are three 
suggestions I have made and there is some similar ideas posted by others today.

1) Perhaps the young rapidly growing trees slow down their growth rate or net 
growth may actually stop or decrease as their top is battered by weather and 
other elements.  Bob Leverett said something similar this morning "Several 
years ago, Will and I begin talking about the possibility that many species 
appear to reach their greatest heights earlier than we would have thought and 
then stagnate or lose height in advancing age.  Tuliptrees in the Smokies fit 
that profile well."

2) The second idea was that these early fast growing tall trees were over time 
replaced by longer lived, but shorter specimens over time resulting in the 
lowering of the net canopy height in subsequent generations.  The idea 
essentially stated in another way is that the tallest stage of the forest is a 
passing phase in the sequence of forest development, but that a shorter forest, 
perhaps with longer lived trees, would form the canopy of a longer term more 
stable forest regime.  We know that some of the oldest specimens for a species 
are often those stunted specimens clinging for life on a rocky cliff somewhere 
so maybe somewhat shorter trees would come to dominate a forest over time.  I 
like this idea conceptually, but have no specific data that would either 
support or oppose it.

3) The third idea was that the circumstances of how the growth of the grove was 
initiated would play a role.  There might be a difference in species 
composition and competition between those groves started from smaller openings 
within a forested setting, and those formed from a large scale disturbance such 
as a major fire or large scale blow down.  The part left out of this idea was 
that the first generation after a large scale reset perhaps could grow taller 
than subsequent generations of replacement trees.  This is similar to the 
second idea above, and both could work together.  The difference is that in the 
second idea the second generation shorter trees may reflect an adaptive genetic 
component favoring longer lived trees, while the third idea postulates that 
different initial growing conditions may be reflected in different heights of 
trees.  Again this is just speculation on how things might work, rather than 
interpretation of data sets.

I an case, whether the ideas are valid or not, they do provide a different 
approach to the question.

Ed Frank
  
http://nature-web-network.blogspot.com/
http://primalforests.ning.com/
http://www.facebook.com/profile.php?ref=profile&id=709156957
  ----- Original Message ----- 
  From: Edward Frank 
  To: [email protected] 
  Sent: Tuesday, January 12, 2010 2:40 AM
  Subject: Re: [ENTS] Re: Projected heights


  Josh, Jess, Will, Gaines, Lee, ENTS,

  You are reporting these taller younger forests.  I am wondering if these 
forests maintain their height or grow taller after this initial youthful 
exuberance of growth?  Could it be possible that the upward growth slows or or 
stops and the trees over the course of time get beaten back down to lower 
levels by 100- 200 year storm events?  Upward growth would not actually cease 
but if it fell below the rate of height loss from weather, the net result would 
be a decrease in the average height of the forest. Perhaps the taller younger 
trees tend to die off after a period of time and are replaced by longer lived, 
but shorter specimens of the same species.  You know that slow growing stunted 
trees clinging to cliffs under poor conditions are often among the oldest for 
the species.  This may not be the case for the forests you are observing, but 
it is an alternative to the idea that environmentally something must have 
changed, and certainly is an alternative to the idea that  indigenous cultures 
somehow did something on a grand scale for which there is little or no evidence 
that made these original "old growth forests" be shorter than younger modern 
forests.  The idea essentially stated in another way is that the tallest stage 
of the forest is a passing phase in the sequence of forest development, but 
that a shorter forest, perhaps with longer lived trees, would form the canopy 
of a longer term more stable forest regime.  

  Another idea might be that the type of regrowth might play a role - the 
height of tree the forest in might reflect whether or not they grew from canopy 
gaps in a broad forest, or from a larger scale disturbance event - like a major 
fire or large area blow down.  These could have different species recruitment, 
and certainly different competition between trees as they grew.  I would think 
it might have an impact on the total height of the resulting forest.  Perhaps 
both postulated mechanisms could play a role.  I think at least the first idea 
has some merit and deserves consideration, it might not be correct, but gives 
another approach to the problem that may be useful even if wrong.
  Ed Frank  

  http://nature-web-network.blogspot.com/
  http://primalforests.ning.com/
  http://www.facebook.com/profile.php?ref=profile&id=709156957

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