Hi, Russell.

Across taxa/clades, sure: clads (Maliet et al., 2019, 
https://doi.org/10.1038/s41559-019-0908-0) or MiSSE (Vasconcelos et al., 2022, 
https://doi.org/10.1111/evo.14517) can do this. They paint different 
diversification rates on different parts of the tree without requiring a priori 
assignment.

However, I’d be careful about interpreting rates through time, even with these 
models. Louca & Pennell (2020, 
https://www.nature.com/articles/s41586-020-2176-1) show that multiple different 
rates can be equally likely. You could use clads or MiSSE to paint ancestral 
nodes into diversification regimes, but I’d worry about how much they actually 
mean.

Best,
Brian

_________________________________________

Brian O’Meara
He/Him
Professor, Dept. of Ecology & Evolutionary Biology
University of Tennessee, Knoxville


From: R-sig-phylo <[email protected]> on behalf of Russell 
Engelman <[email protected]>
Date: Tuesday, July 23, 2024 at 8:42 AM
To: mailman, r-sig-phylo <[email protected]>
Subject: [R-sig-phylo] Calculating Asymmetry in Speciation Across a Phylogeny
Dear R-Sig-Phylo,

I was wondering if there was a way to quantitatively examine
diversification and speciation rates that is sensitive to the fact that not
all clades diversify equally?

I have a phylogeny where the overall diversity of the group — particularly
in terms of higher level clades — declines at a specific point in time, but
the broader pattern is being covered up by a massive burst of speciation at
one ecologically stereotyped (and possibly taxonomically oversplit) node on
the tree. Raw speciation and extinction rates end up giving a biased view
of the pattern because they treat all lineages as equal and can't parse
that 6-10 clades are declining while the remaining one is undergoing
diversification. This contrasts with older parts of the tree where
diversification is more evenly spread across different clades.

I had considered calculating a Shannon Diversity Index or Inverse Simpson's
Diversity Index at each point on a LTT after dividing the tree up by family
to get an idea of the overall balance of alpha diversity across clades over
time (i.e., clade X represents a disproportionate contribution to standing
alpha diversity as time goes on). However, this relies on dividing the tree
into a priori groups based on relatively arbitrary criteria and felt rather
hacky. I was wondering if there was a more impartial way of calculating or
quantitatively describing the asymmetry in diversification rates across
nodes over time?

Sincerely,
Russell

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