My understanding of wheel taper (And I may be full of B.S., but this is  what 
I have read) is to allow trains to navigate curves without excessive wheel  
slip. As centrifugal force causes the wheel set to move to the outside of  the 
curve the larger diameter of the tread is contacting the rail  on the larger 
diameter, while the inner wheel is riding on its smaller diameter.  This 
lessens wheel slip and I presume that on 1=1 scale (full size) that  railroads 
set 
minimum curve radius to suit on mainline practice. Look at the  steep taper on 
a Lionel 027 wheel compared to the slight taper on a fine scale  wheel. 
 
I have two kinds of wheels on my 3.5 gauge stuff, one set has each wheel  
with a brass bushing so the wheels can rotate independent of what his brother 
on  
the other end of the axle is doing. The other sets of wheels each wheel is 
press  fitted on the axle so where "Righty" goes, "Lefty" must follow along. 
Next time  I am out at the shop will check and see if either, both or which 
ever 
has taper  and which don't. I have seen no difference in the wheel sets to 
stay on the  track over my 20' radius curves. 
 
Of course on a rod driven locomotive the wheels must be secured to each  axle 
to keep the valve gear working properly. 
 
On my lathe (Sear's Atlas, 12 inch) I have no problem turning the slight 3  
degree taper by setting over the compound to three degrees. Or it could be done 
 by a file on our small wheels with a template to see where we are in the  
process. The wheels must be made to the same diameter, coned or flat within as  
close a limit as you can manage so both sides work the same. What I have done  
and seen recommended is to "rough" turn all your wheels first, then set up to 
 finish each wheel at the same settings on your cross slide and compound. 
When I  have done cast iron wheels I use a carbide bit to dig under the cast 
iron 
"skin"  then when all wheels in the set are rough turned leaving a few thous 
for  finishing change to a HSS tool with the root radius between tread and 
flange  ground in. Then set up a mandrel to take each wheel running true on its 
axis and  take off the final few thous and put in the taper and touch the outer 
rim with a  file to just barely round it over. Also use a file to radius the 
top of the  flange, but I don't think that is too important, but I presume a 
special ground  tool or jig would do that more evenly. 
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