Dear all,

to trim or not to to trim….

- what is the difference between highly disordered side-chain atoms versus 
disordered bulk-solvent atoms? Do not both represent a continuum function with 
a comparable electron-density distribution? Have not solvent and side-chain 
atoms merged/mixed at this position in space and with this, are effectively 
indistinguishable (contrast loss i.e. similar contribution to diffraction) from 
one another?

- are current solvent models comprehensive enough in that modelling of atoms 
with very high B factors is really mutually beneficial for bulk and structure 
model?

- is model bias (overfitting) not increased by modelling highly disordered 
side-chain atoms (Rfree will probably not increase but R will drop)?

- analogous to the disordered side-chain atoms, bulk solvent atoms are of 
course also present in the crystal, so why not fill the AU with solvent 
molecules and let refinement take care of them?
(published methodology, see https://doi.org/10.1107/S0907444906021627 
<https://doi.org/10.1107/S0907444906021627>)


Trimming or non-evidence-based placement and refinement are somehow halfhearted 
options with split opinions among the community as the discussions demonstrate.

Adding/Modelling a plausible side-chain rotamer with zero occupancy still has 
its charm and better reflects the situation "the side-chain atoms must be there 
but we do not know exactly where"……..

Cheers

Jeroen



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